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Linux and OSX clients will not use the GUP functionality at all.
Depending on the mixture of operating systems within your environment, this is important to know.
In this scenario none of the agents will actually communicate with another GUP since a GUP can only retrieve updates from a SEPM.
This is a scenario where some people could believe that the environment would act in a full peer-to-peer fashion.
If the GUP does not have a definition it will reach out to its defined SEP Manager and download the correct update.
Since the controls are in place to insure that definitions originate from a SEPM to the GUP this will not occur.
Things you need to be aware of when using Group Update Providers The most important thing to understand is that GUPs only work with Windows clients.
A few months later it was discovered that these systems have been decommissioned and replaced.
During this time while they thought a GUP was being used, all clients at this remote site were actually updating definition directly from the SEPM.